The Penguins of Madagascar is a spin-off from the film Madagascar. The film series follows the adventures of four penguin protagonists: Skipper, Kowalski, Rico, and Private, who carry out paramilitary missions such as the mission to protect their homes in the Central Park Zoo. Penguins often deal with problems caused or exacerbated, by King Julien XIII (ring tail lemurs), Maurice (an aye-aye), and Mort (rat lemurs).
It is not known how penguins and lemurs arrive at the zoo after their adventures in both Madagascar films. During the opening title of the event, penguins were also seen opening a chest that read "Madagascar" and contained three lemur characters. DreamWorks Animation CEO Jeffrey Katzenberg stated that "there is at least one more chapter. We finally want to see those characters returning to New York."
Skipper Coloring Pages For Kids (Tom McGrath) is the leader of the penguins; he designed tactics and gave orders. Count and can hardly be disturbed. Paranoia Skipper is delirious and the tendency to see everything as a military operation, combined with his experience in secret forces, has encouraged him to prepare himself in any situation, no matter how strange or not.
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Private (James Patrick Stuart), is an emotional and sensitive new warrior in the penguin group. Although younger and less experienced than other penguins, he is the most humble. Private tends to offer simpler, more sensible solutions in response to Kowalski's complex strategy for Skipper.
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Rico (John DiMaggio), is a weapons team and explosives specialist, who primarily communicates through growls and screams. Slightly flexed, Rico swallowed useful tools, such as dynamite, and spit it out when needed.
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King Julien XIII (Danny Jacobs), usually abbreviated as King Julien, is a lover of pleasure, and the ring-tailed slow loris. The lemur king is a funny and arrogant character who has little attention for others, even his people. He has a tendency to use malapropisms and misinterpret in his figurative speech.
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Maurice (Kevin Michael Richardson), an aye-aye, is one of the people of King Jullien. Maurice accepted his life as a servant, but he often showed disdain for King Julien and his indifferent attitude.
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Mort (Andy Richter), is a vigorous slow loris and a rat loris that is prone to accidents. Unlike Maurice, Mort showed his loyalty to the king of Julien, even showing his obsession at the feet of King Julien. But King Julien treated him with contempt.
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Marlene (Nicole Sullivan), is a female beaver who is transferred to the Central Park Zoo from an aquarium in California. He was a close friend of Skipper and sometimes trailed Skipper and other penguins on missions, but he was an impartial character between penguins and lemurs. With a cool head, and a feminine voice, he is a rival to Skipper's masculine character, a secret troop character. Because Marlene was born in captivity, when she crossed the zoo boundary, she went berserk.
Mason (Conrad Vernon) and Phil, are two intelligent chimpanzees. Mason can talk but cannot read, Phil is silent but loyal reader and communicates through sign language interpreted by Mason.